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In a study by Hosseini et al

They are produced intracellularly in several stages: stages I and II include unmelanized immature premelanosomes in the middle of the melanocyte, whereas stages III and IV contain melanized melanosomes migrating to the outer edge of the cell through the dendrites 31

Polymorphism of GSTA1 is represented by three linked single nucleotide polymorphisms (SNPs), resulting in differential expression with lower transcriptional activation of the variant GSTA1 * B (T) than the common GSTA1*A (C) allele 33