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APS improve ER and the resulting autophagy by inhibiting the ATF6 passage and glucose-regulated protein 78 (GRP78)-related pathways [including IRE-1, the c-Jun N-terminal kinase (JNK), and PERK], reducing the apoptosis factors expression of C/EBP-homologous protein and caspase-12 (Wang et al., 2009

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The histidine imidazole ring, glycine amino groups, and deprotonated amide nitrogens coordinate the copper ion, creating a stable chelation complex